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How does FastQTL deal with strand

FastQTL phenotype.bed input does not have strand column as in QTLtools. How does the software deal with strand? I guess a related question would be say gene1 is on the reverse/minus strand in GTF file. Should its end position in GTF file be the TSS (transcription start site) for fastQTL phenotype.bed input? Thanks in advance.

rna-seq qtl

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