I can't seem to find metadata on the output of GeneMarkS-2. I'm trying to figure out what the last 4 columns mean in the data below in particular. Any guidance would be greatly appreciated. Thank you.

# GeneMark.hmm-2 version: 1.04

# File with sequence: seq.fna

# File with native parameters: GMS2.mod

# File with MetaGeneMark parameters: /home/genemark/bin/gms2/mgm_11.mod

#

# start: Sat Oct 26 22:01:09 2019

# Sequence meta data: >CP033686.1 Pseudomonas aeruginosa strain H25883 chromosome, complete genome

# sequence-region 1 6706800

SequenceID: 1

     1   +    22    1539    1518 native TTGGCA 7 1
     2   +    1568    2671    1104 native AGGGAC 6 1
     3   +    2681    3790    1110 atypical AATCAC 5 1