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nucleotide Hidden Markov Model search

I am using HMMER v3.1 for nucleotide Hidden Markov Model searches.

What I can't find in the manual is whether the hmm (I am building species-specific domain specific hmm) used to search a database looks at the plus and minus strand of the sequences in the database. To clarify - when running a BLAST the output gives matches against both forward and reverse. I am not sure if this occurs with HMMER. Can anyone help with this?

nhmmer hmmer

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